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Kingman Coalescent Tree Moments

Build genealogy moments from independent merger waits and explain mutation-count overdispersion.

Free theory article

Build genealogy moments from independent merger waits and explain mutation-count overdispersion.

Concept & interpretation

The Kingman Coalescent models the backward-in-time genealogy of a sample of n lineages, assuming a neutrally evolving constant-size population. Pairs of lineages merge uniformly at random, with standard waiting times driven by independent Exponential distributions.

Experiment

You can interactively explore these properties in the module at https://riano.app/module/kingman-coalescent-tree-moments.

  1. Select Sample lineages to 6. Observe that the tree gains more stages and the Expected total branch length increases to approximately 4.566666666666666.
  2. Change the Mutation theta to 2. Notice that the Expected segregating sites doubles because mutations fall independently along the branches with rate proportional to theta.
  3. Change the Stage probe to 3. This highlights the tree where 3 ancestral lineages remain. The Merger Rate becomes 3.0 and the Mean Wait becomes 0.3333333333333333.
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